Platanus hispanica

Platanus × acerifolia, the London plane, London planetree, or hybrid plane, is a tree in the genus Platanus. It is often known by the synonym Platanus × hispanica. It is usually thought to be a hybrid of Platanus orientalis (oriental plane) and Platanus occidentalis (American sycamore). Some authorities think that it may be a cultivar of P. orientalis. The London plane is a large deciduous tree growing 20–30 m (66–98 ft), exceptionally over 40 m (131 ft) tall, with a trunk up to 3 m (10 ft) or more in circumference. The bark is usually pale grey-green, smooth and exfoliating, or buff-brown and not exfoliating. The leaves are thick and stiff-textured, broad, palmately lobed, superficially maple-like, the leaf blade 10–20 cm (4–8 in) long and 12–25 cm (5–10 in) broad, with a petiole 3–10 cm (1–4 in) long. The young leaves in spring are coated with minute, fine, stiff hairs at first, but these wear off and by late summer the leaves are hairless or nearly so. The flowers are borne in one to three (most often two) dense spherical inflorescences on a pendulous stem, with male and female flowers on separate stems. The fruit matures in about 6 months, to 2–3 cm diameter, and comprises a dense spherical cluster of achenes with numerous stiff hairs which aid wind dispersal; the cluster breaks up slowly over the winter to release the numerous 2–3 mm seeds. The London Plane is one of the most efficient trees in removing small particulate pollutants in urban areas. It shares many visual similarities with Platanus occidentalis (American sycamore), from which it is derived; however, the two species are relatively easy to distinguish, considering the London plane is almost exclusively planted in urban habitats, while P. occidentalis is most commonly found growing in lowlands and alluvial soils along streams. The species was formed by hybridization in the 17th century after P. orientalis and P. occidentalis had been planted in proximity to one another. It is often said that the hybridization took place in Spain, but it could also have happened in Vauxhall Gardens in London where John Tradescant the Younger discovered the tree in the mid-17th century.The leaf and flower characteristics are intermediate between the two parent species, the leaf being more deeply lobed than P. occidentalis but less so than P. orientalis, and the seed balls typically two per stem (one in P. occidentalis, 3–6 in P. orientalis). The hybrid is fertile, and seedlings are occasionally found near mature trees. Controlled reciprocal pollinations between P. occidentalis and P. orientalis resulted in good yields of germinable seed and true hybrid seedlings. Crosses of both species, as females, with P. racemosa and P. wrightii produced extremely low yields of germinable seed, but true hybrids were obtained from all interspecific combinations. Apomixis (asexual reproduction from non-fertilized seeds) appeared common in P. orientalis. In 1968 and 1970, Frank S. Santamour Jr. recreated the P. orientalis by P. occidentalis cross using a P. orientalis of Turkish origin with American sycamores (P. occidentalis). The offspring were evaluated following several years of exposure to anthracnose infection. Two selections, 'Columbia' and 'Liberty', were released in August, 1984. Platanus × acerifolia was first formally described in the botanical literature by the Scottish botanist William Aiton in his 1789 work Hortus Kewensis as a variety of P. orientalis. Aiton described this variety with a two-word Latin diagnosis, 'foliis transversis', and called it the Spanish plane tree. In 1805, Carl Ludwig Willdenow chose to elevate Aiton's variety to species rank, publishing the new species P. acerifolia in the fourth edition of Species Plantarum. The species name was then modified to include the multiplication symbol to indicate its suspected hybrid parentage. The other name commonly used for this taxon, Platanus × hispanica auct. non Mill. ex Münchh., is a nomen dubium based on an uncertain description. The London plane is one of 50 Great British Trees The Tree Council selected in 2002 in honour of Queen Elizabeth II's Golden Jubilee. The list specifically mentions Britain's first London plane being in the city of Ely, Cambridgeshire.