Coral

Corals are marine invertebrates within the class Anthozoa of the phylum Cnidaria. They typically live in compact colonies of many identical individual polyps. Corals species include the important reef builders that inhabit tropical oceans and secrete calcium carbonate to form a hard skeleton.Timeline of the major coral fossil record and developments from 650 m.y.a. to present.Fungia sp. skeletonPolyps of Eusmilia fastigiataPillar coral, Dendrogyra cylindricusBrain coral, Diploria labyrinthiformisBrain coral spawningBrain coral releasing eggsFringing coral reef off the coast of Eilat, Israel. Corals are marine invertebrates within the class Anthozoa of the phylum Cnidaria. They typically live in compact colonies of many identical individual polyps. Corals species include the important reef builders that inhabit tropical oceans and secrete calcium carbonate to form a hard skeleton. A coral 'group' is a colony of myriad genetically identical polyps. Each polyp is a sac-like animal typically only a few millimeters in diameter and a few centimeters in length. A set of tentacles surround a central mouth opening. An exoskeleton is excreted near the base. Over many generations, the colony thus creates a large skeleton characteristic of the species. Individual heads grow by asexual reproduction of polyps. Corals also breed sexually by spawning: polyps of the same species release gametes simultaneously over a period of one to several nights around a full moon. Although some corals are able to catch small fish and plankton using stinging cells on their tentacles, most corals obtain the majority of their energy and nutrients from photosynthetic unicellular dinoflagellates in the genus Symbiodinium that live within their tissues. These are commonly known as zooxanthellae. Such corals require sunlight and grow in clear, shallow water, typically at depths less than 60 metres (200 ft). Corals are major contributors to the physical structure of the coral reefs that develop in tropical and subtropical waters, such as the enormous Great Barrier Reef off the coast of Queensland, Australia. Other corals do not rely on zooxanthellae and can live in much deeper water, with the cold-water genus Lophelia surviving as deep as 3,300 metres (10,800 ft). Some have been found as far north as the Darwin Mounds, northwest of Cape Wrath, Scotland, and others off the coast of Washington State and the Aleutian Islands. Aristotle's pupil Theophrastus described the red coral, korallion, in his book on stones, implying it was a mineral, but he described it as a deep-sea plant in his Enquiries on Plants, where he also mentions large stony plants that reveal bright flowers when under water in the Gulf of Heroes. Pliny the Elder stated boldly that several sea creatures including sea nettles and sponges 'are neither animals nor plants, but are possessed of a third nature (tertius natura)'. Petrus Gyllius copied Pliny, introducing the term zoophyta for this third group in his 1535 book On the French and Latin Names of the Fishes of the Marseilles Region; it is popularly but wrongly supposed that Aristotle created the term. Gyllius further noted, following Aristotle, how hard it was to define what was a plant and what was an animal. The Persian polymath Al-Biruni (d. 1048) classified sponges and corals as animals, arguing that they respond to touch. Nevertheless, people believed corals to be plants until the eighteenth century, when William Herschel used a microscope to establish that coral had the characteristic thin cell membranes of an animal. Presently, corals are classified as certain species of animals within the sub-classes Hexacorallia and Octocorallia of the class Anthozoa in the phylum Cnidaria. Hexacorallia includes the stony corals and these groups have polyps that generally have a 6-fold symmetry. Octocorallia includes blue coral and soft corals and species of Octocorallia have polyps with an eightfold symmetry, each polyp having eight tentacles and eight mesenteries. Fire corals are not true corals, being in the order Anthomedusae (sometimes known as Anthoathecata) of the class Hydrozoa. Corals are sessile animals and differ from most other cnidarians in not having a medusa stage in their life cycle. The body unit of the animal is a polyp. Most corals are colonial, the initial polyp budding to produce another and the colony gradually developing from this small start. In stony corals, also known as hard corals, the polyps produce a skeleton composed of calcium carbonate to strengthen and protect the organism. This is deposited by the polyps and by the coenosarc, the living tissue that connects them. The polyps sit in cup-shaped depressions in the skeleton known as corallites. Colonies of stony coral are very variable in appearance; a single species may adopt an encrusting, plate-like, bushy, columnar or massive solid structure, the various forms often being linked to different types of habitat, with variations in light level and water movement being significant.