Asiatic wild dog

The dhole /doʊl/ (Cuon alpinus) is a canid native to Central, South, and Southeast Asia. Other English names for the species include Asian wild dog, Asiatic wild dog, Indian wild dog, whistling dog, red dog, and mountain wolf. It is genetically close to species within the genus Canis,(Fig. 10) but distinct in several anatomical aspects: its skull is convex rather than concave in profile, it lacks a third lower molar and the upper molars sport only a single cusp as opposed to between two and four. During the Pleistocene, the dhole ranged throughout Asia, Europe, and North America but became restricted to its historical range 12,000–18,000 years ago. The dhole is a highly social animal, living in large clans without rigid dominance hierarchies and containing multiple breeding females. Such clans usually consist of 12 individuals, but groups of over 40 are known. It is a diurnal pack hunter which preferentially targets medium- and large-sized ungulates. In tropical forests, the dhole competes with tiger and leopard, targeting somewhat different prey species, but still with substantial dietary overlap. It is listed as Endangered by the IUCN as populations are decreasing and are estimated at fewer than 2,500 adults. Factors contributing to this decline include habitat loss, loss of prey, competition with other species, persecution due to livestock predation and disease transfer from domestic dogs. The etymology of 'dhole' is unclear. The possible earliest written use of the word in English occurred in 1808 by soldier Thomas Williamson, who encountered the animal in Ramghur district. He stated that dhole was a common local name for the species. In 1827, Charles Hamilton Smith claimed that it was derived from a language spoken in 'various parts of the East'. Two years later, Smith connected this word with Turkish: deli 'mad, crazy', and erroneously compared the Turkish word with Old Saxon: dol and Dutch: dol (cfr. also English: dull; German: toll), which are in fact from the Proto-Germanic *dwalaz 'foolish, stupid'. Richard Lydekker wrote nearly 80 years later that the word was not used by the natives living within the species' range. The Merriam-Webster Dictionary theorises that it may have come from the Kannada: tōḷa ('wolf'). Canis alpinus was the binomial name proposed by Peter Simon Pallas in 1811, who described its range as encompassing the upper levels of Udskoi Ostrog in Amurland, towards the eastern side and in the region of the upper Lena River, around the Yenisei River and occasionally crossing into China. This northern Russian range reported by Pallas during the 18th and 19th centuries is 'considerably north' of where this species occurs today. Canis primaevus was a name proposed by Brian Houghton Hodgson in 1833 who thought that the dhole is a primitive Canis form and the progenitor of the domestic dog. Hodgson later took note of the dhole's physical distinctiveness from the genus Canis and proposed the genus Cuon. The first study on the origins of the species was conducted by paleontologist Erich Thenius, who concluded that the dhole was a post-Pleistocene descendant of a golden jackal-like ancestor. The earliest known member of the genus Cuon is the Chinese Cuon majori of the Villafranchian period. It resembled Canis in its physical form more than the modern species, which has greatly reduced molars, whose cusps have developed into sharply trenchant points. By the Middle Pleistocene, C. majori had lost the last lower molar altogether. C. alpinus itself arose during the late Middle Pleistocene, by which point the transformation of the lower molar into a single cusped, slicing tooth had been completed.