Populus trichocarpa

Populus trichocarpa, the black cottonwood, western balsam-poplar or California poplar, is a deciduous broadleaf tree species native to western North America. It is used for timber, and is notable as a model organism in plant biology. A high-coverage genome sequence was published in 2006 and was the first tree genome to be sequenced. Long-read Pacbio sequencing enabled the assembly of all scaffolds in 2018. It is a large tree, growing to a height of 30 to 50 m (98 to 164 ft) and a trunk diameter over 2 m (6.6 ft), which makes it the largest poplar species in the Americas. It is normally fairly short-lived, but some trees may live up to 400 years. A cottonwood in Willamette Mission State Park near Salem, Oregon, holds the national and world records. Last measured in April, 2008, this black cottonwood was found to be standing at 155 ft (47 m) tall, 29 ft (8.8 m) around, with 527 points. The bark is grey and covered with lenticels, becoming thick and deeply fissured on old trees. The bark can become hard enough to cause sparks when cut with a chainsaw. The stem is grey in the older parts and light brown in younger parts. The crown is usually roughly conical and quite dense. In large trees, the lower branches droop downwards. Spur shoots are common. The wood has a light coloring and a straight grain. The leaves are 7–20 cm long with a glossy, dark green upper side and glaucous, light grey-green underside; larger leaves, up to 30 cm long, may be produced on stump sprouts and very vigorous young trees. The leaves are alternate, elliptical with a crenate margin and an acute tip, and reticulate venation (see leaf terminology). The petiole is reddish. The buds are conical, long, narrow, and sticky, with a strong balsam scent in spring when they open. P. trichocarpa has an extensive and aggressive root system, which can invade and damage drainage systems. Sometimes, the roots can even damage the foundations of buildings by drying out the soil. Flowering and fruiting P. trichocarpa is normally dioecious; male and female catkins are borne on separate trees. The species reaches flowering age around 10 years. Flowers may appear in early March to late May in Washington and Oregon, and sometimes as late as mid-June in northern and interior British Columbia, Idaho, and Montana. Staminate catkins contain 30 to 60 stamens, elongated to 2 to 3 cm, and are deciduous. Pistillate catkins at maturity are 8 to 20 cm long with rotund-ovate, tricarpellate subsessile fruits 5 to 8 mm long. Each capsule contains many minute seeds with long, white, cottony hairs. Seed production and dissemination