In Memoriam: David L. Nanney (1925–2016): Tetrahymena Genetics Founder and Epigenetics Champion

![David L. Nanney][1] David L. Nanney David Ledbetter Nanney, who passed away on June 4, 2016, was a distinguished geneticist, cell, and developmental biologist who founded Tetrahymena genetics and reinvented the word “epigenetics.” Nanney was truly a natural historian, with a gift for