A revision of the Phelister haemorrhous species group (Coleoptera, Histeridae, Exosternini)

The Phelisterhaemorrhous species group is established here, revising the seventeen included species, four of which are described as new. This group is named for and contains the type species of Phelister, so represents a core around which a modern concept of the dumping-ground genus Phelister may be developed. The group includes several common and well-known species in the Americas, including some of the only Phelister to exhibit distinctive coloration. Several of these are typically found in cattle dung, and have likely expanded beyond their native ranges as cattle spread throughout the Americas. The group contains the following species: Phelisterhaemorrhous Marseul, 1854, Phelisteraffinis J.E. LeConte, 1859, Phelisterparallelisternus Schmidt, 1893, Phelistermobilensis Casey, 1916, Phelisterbrevistriatus Casey, 1916, Phelistersonoraesp. nov., Phelisterwarnerisp. nov., Phelisterpuncticollis Hinton, 1935, Phelistersubrotundus (Say, 1825), Phelisterrouzeti (Fairmaire, 1850), Phelisterrufinotus Marseul, 1861, Phelisterthiemei Schmidt, 1889, Phelisterparecissp. nov., Phelisterbryantisp. nov., Phelistervernus (Say, 1825), Phelisterchilicola Marseul, 1870, and Phelisterbruchi Bickhardt, 1920. We also designate the following new synonymies: Phelisterhaemorrhous Marseul (= Phelisterrubicundus Marseul, 1889, syn. nov.); Phelistersubrotundus (= Phelistercontractus Casey, 1916, syn nov.); Phelisterrouzeti (Fairmaire) (= Phelisterfairmairei Marseul, 1861; syn. nov., = Phelisterwickhami Casey, 1916, syn. nov.); Phelisterrufinotus Marseul, 1861 (= Epierusmarseulii Kirsch, 1873, syn. nov.); and Phelisterthiemei Schmidt, 1889 (= Phelisterstercoricola Bickhardt, 1909, syn. nov.).