Sphaerapus, A Poorly Known Invertebrate Trace Fossil From Nonmarine Permian and Jurassic Strata of North America

In 1858, Edward Hitchcock named Sphaerapus larvalis and S. magnus for burrows from the nonmarine Lower Jurassic Turners Falls Formation in Massachusetts. Walpia hermitensis White, 1929, from the Lower Permian Hermit Shale, Arizona, is also a little known ichnotaxon. Examination of the type material indicates that Sphaerapus is a senior subjective synonym of Walpia. Sphaerapus is also a nomen oblitum, but Walpia has not been sufficiently used to replace its senior synonym as a conserved name; thus, we use the senior synonym Sphaerapus as the valid name of this ichnogenus, which is monospecific (S. larvalis = S. magnus = W. hermitensis). Numerous specimens of Sphaerapus from the Lower Permian of New Mexico and Texas, and a new record from the Lower Jurassic of Massachusetts show that it is a horizontal endichnion or exichnion having an unpacked tunnel with a cylindrical to ellipsoidal cross section, in some specimens branching, and lined with flattened, lenticular, non-coprolitic sediment pellets that are i...