Parahiraciini (Hemiptera, Fulgoromorpha, Issidae): one new genus, two new species and three new subtribes

A new genus Pusulissus gen. nov. with two new species P. phiaoacensis sp. nov. and P. coronomensis sp. nov. are described respectively from Vietnam and China in the tribe Parahiraciini of the family Issidae. A molecular phylogeny using combined genes (18S, 28S, COX1 and Cytb) confirms its placement in the tribe Parahiraciini. The tribe is divided into three subtribes: Scantiniina subtribe nov. with the type genus Scantinius Stal, 1866, and Vindilisina subtribe nov. with type genus Vindilis Stal, 1870 plus Nisoprincessa Gnezdilov, 2017, and the nominal subtribe Parahiraciina Cheng & Yang, 1991 subtribe nov. The characteristics of these subtribes are provided, with a key to identification. Genus Folifemurum Che, Zhang & Wang, 2013 is transferred to Hemisphaeriini Mongolianina, and genus Gelastyrella Yang, 1994 is maintained as a valid name.