A New Primitive Therocephalian (Theromorpha) from the Middle Permian of Eastern Europe

The largest East European scylacosaurid therocephalian Julognathus crudelis gen. et sp. nov. is described based on materials from the middle Permian locality of Sundyr-1, Mari El Republic, Russia. Julognathus occupied the niche of the largest predator within the Sundyr tetrapod community, forming a trophic match with the herbivorous tapinocephalian.