First record of the pterosaur footprint Pteraichnus from the Saltwick Formation (Aalenian) of the Cleveland Basin, Yorkshire, UK

Summary Prints belonging to Pteraichnus are recorded and described from the Saltwick Formation (Middle Jurassic, Aalenian Stage) at Hayburn Wyke on the Cleveland Coast of Yorkshire, England. This is the first record of Pteraichnus from the region and from the British Isles, and is an important addition to its diverse vertebrate ichnofauna. It is also only the second record of pterosaur footprints from the British Isles and is the oldest known occurrence of Pteraichnus . Significantly, it suggests that the pterodactyloid pterosaurs evolved earlier than has previously been supposed.