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[Jiang, Lei; Huang, Hui; Yuan, Xiang-Cheng; Yuan, Tao; Zhang, Yu-Yang; Wen, Colin Kuo-Chang; Li, Xiu-Bao; Zhou, Guo-Wei] Chinese Acad Sci, South China Sea Inst Oceanol, Guangdong Prov Key Lab Appl Marine Biol, Key Lab Trop Marine Bioresources & Ecol, Guangzhou 510301, Guangdong, Peoples R China; [Jiang, Lei; Huang, Hui; Zhou, Guo-Wei] Chinese Acad Sci, Trop Marine Biol Res Stn Hainan, Sanya 572000, Peoples R China; [Jiang, Lei; Zhang, Yu-Yang] Univ Chinese Acad Sci, Beijing 100049, Peoples R China