A review of the Bembidion (Odontium) aenulum subgroup (Coleoptera: Carabidae), with description of a new species

The Bembidion aenulum subgroup of the subgenus Odontium is reviewed. This subgroup, widespread in the United States east of the Rocky Mountains, previously has been considered to consist of only one species, Bembidion aenulum Hayward. However, morphological studies coupled with analyses of 28S ribosomal RNA, cytochrome oxidase I, CAD, and wingless genes reveal that eastern members of this group belong to a new species herein named Bembidion paraenulum new species. This species ranges from New Hampshire in the northeast, through Virginia and North Carolina, south to Florida and west to Mississippi. Both species are described and illustrated.